Grubeosyllis and Exogone (Exogoninae, Syllidae, Polychaeta) from Cuba, the Gulf of Mexico, Florida and Puerto Rico, with a Revision of Exogone

Twenty-one species of the genera Grubeosyllis Verrill, 1900, and Exogone Örsted, 1845 (Exogoninae: Syllidae), principally from Cuba but also from the Gulf of Mexico, Florida and Puerto Rico are reported in this paper. The genus Grubeosyllis is re-erected to replace Pseudobrania San Martín, 1984, a junior synonym. The genus Exogone is revised, three subgenera are proposed: Exogone Örsted, 1845, Parexogone Mesnil and Caullery, 1916, and Sylline Claparède, 1864. Keys of each subgenus for the identification of all species described are given. The following new species are described: Exogone (Parexogone) caribensis, E. (P.) wolfi, Exogone (Exogone) rolani, E. (E.) longispinulata, and E. (E.) pseudolourei. The species and subspecies Grubeosyllis heterocirra (Rioja, 1941); G. vieitezi (San Martín, 1984); G. euritmica (Sardá, 1984); Exogone (Parexogone) exmouthensis Hartmann-Schröder, 1980; Exogone (Parexogone) parahomoseta mediterranea San Martín, 1984; Exogone (Exogone) breviantennata Hartmann-Schröder, 1959; and Exogone (Sylline) naidinoides Westheide, 1974, are new to the Caribbean and Gulf of Mexico area. The species Grubeosyllis nitidula Verrill, 1900; Grubeosyllis rugulosa Verrill, 1900; Exogone (Parexogone) atlantica Perkins, 1981; Exogone (Exogone) dispar (Webster, 1879); E. (E.) lourei Berkeley and Berkeley, 1948; and E. (E.) arenosa Perkins, 1981, are new to the Cuban fauna. The species Grubeosyllis nitidula and G. rugulosa are identified tentatively because there is doubt about their correct determination, and G. nitidula is re-described on the bases of Cuban specimens. Two species of Exogone (Exogone) have been identified only to subgenus.