Phylogeny and systematics of the Arrupinae (Chilopoda Geophilomorpha Mecistocephalidae) with the description of a new dwarfed species

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We performed a cladistic analysis of the subfamily Arrupinae, based on morphological data (23 characters) from all the 16 species we recognize in the group. Based on this analysis, the following taxonomic and nomenclatural novelties are introduced: Nannarrup hoffmani n. g. n. sp. is described from introduced specimens of unknown origin found living in New York, USA; Agnostrup n. g. (type species Krateraspis striganovae Titova, 1975) is created to include A. striatus (Takakuwa, 1949) (from Taiwanella), A. paucipes (Miyosi, 1955) (from Taiwanella ) and A. striganovae (Titova, 1975) (from Krateraspis), all new combinations; Nodocephalus areolatus Shinohara, 1957, N. asiaticus Titova, 1975 and Prolamnonyx mamaevi Titova, 1975 are moved to the genus Arrup (three more new combinations); Partygarrupius Verhoeff, 1939, formerly a subgenus of Tygarrup Chamberlin, 1914, is raised to the status of genus. The peculiar morphology of Nannarrup hoffmani is analysed as a possible instance of miniaturization.

Keywords: Agnostrup; Arrupinae; Chilopoda; Geophilomorpha; Mecistocephalidae; Nannarrup; cladistics; miniaturization; taxonomy

Document Type: Research Article


Affiliations: 1: Dipartimento di Biologia, Università degli Studi di Padova, Via Ugo Bassi 58b, I-35131 Padova, Italy; e-mail:,, 2: Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, Paseo del Bosque s.n., (1900) La Plata, R. Argentina; e-mail:

Publication date: October 1, 2003

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