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Morphological Changes in Leaves of Residual Western Hemlock after Clear and Shelterwood Cutting

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The change in microclimate associated with clear and shelterwood cutting had marked effects on the leaves of residual western hemlock trees. In a new clearcutting, old needles abscised prematurely and needles that formed during the first season of exposure were smaller than normal, apparently because of water stress. Needles formed in the second season had thicker palisade mesophyll, a smaller ratio of surface area to weight, and more stomata per unit surface area. Leaves that had developed in the shade and were later exposed showed a slight increase in thickness, probably because of increased palisade mesophyll development. These changes in leaf area and structure were toward a configuration which was more efficient in utilization of light and water in a clearcutting. Hemlock saplings left in a shelterwood suffered less needle loss and mortality, but they still developed needles adapted to full sunlight, suggesting a shelterwood as a beneficial treatment for understory hemlock. Forest Sci. 23:195-203.
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Keywords: Palisade mesophyll; Tsuga heterophylla; microclimate; ratio of surface area to weight; stomatal frequency

Document Type: Journal Article

Affiliations: Research Associate, School of Forestry, Oregon State University, Corvallis, OR 97331

Publication date: 1977-06-01

More about this publication?
  • Forest Science is a peer-reviewed journal publishing fundamental and applied research that explores all aspects of natural and social sciences as they apply to the function and management of the forested ecosystems of the world. Topics include silviculture, forest management, biometrics, economics, entomology & pathology, fire & fuels management, forest ecology, genetics & tree improvement, geospatial technologies, harvesting & utilization, landscape ecology, operations research, forest policy, physiology, recreation, social sciences, soils & hydrology, and wildlife management.
    Forest Science is published bimonthly in February, April, June, August, October, and December.

    2016 Impact Factor: 1.782 (Rank 17/64 in forestry)

    Average time from submission to first decision: 62.5 days*
    June 1, 2016 to Feb. 28, 2017

    Also published by SAF:
    Journal of Forestry
    Other SAF Publications
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