@article {Hillig:March 2005:0925-9864:161, author = "Hillig, Karl", title = "Genetic evidence for speciation in Cannabis (Cannabaceae)", journal = "Genetic Resources and Crop Evolution", volume = "52", year = "March 2005", abstract = "Sample populations of 157 Cannabis accessions of diverse geographic origin were surveyed for allozyme variation at 17 gene loci. The frequencies of 52 alleles were subjected to principal components analysis. A scatter plot revealed two major groups of accessions. The sativa gene pool includes fiber/seed landraces from Europe, Asia Minor, and Central Asia, and ruderal populations from Eastern Europe. The indica gene pool includes fiber/seed landraces from eastern Asia, narrow-leafleted drug strains from southern Asia, Africa, and Latin America, wide-leafleted drug strains from Afghanistan and Pakistan, and feral populations from India and Nepal. A third putative gene pool includes ruderal populations from Central Asia. None of the previous taxonomic concepts that were tested adequately circumscribe the sativa and indica gene pools. A polytypic concept of Cannabis is proposed, which recognizes three species, C. sativa, C. indica and C. ruderalis, and seven putative taxa.", pages = "161-180(20)", url = "http://www.ingentaconnect.com/content/klu/gres/2005/00000052/00000002/00004452" doi = "doi:10.1007/s10722-003-4452-y" }